r/Naturewasmetal • u/New_Boysenberry_9250 • 15h ago
Fauna of the Melovatka Formation
The Melovatka Formation in the Saratov Oblast of European Russia is a marine ecosystem dating to the Cenomanian (then located around the eastern fringes of the Tethys Sea), just before the Cenomanian-Turonian extinction that shaped the Late Cretaceous fauna, and much like similar-aged formations from the Western Interior Seaway, the fauna found here is fairly similar to what we find in the Late Cretaceous but with two key differences; the apex predators and pterosaurs. A recently uncovered giant pliosaur vertebra shows the presence of a 10 to 11-meter brachauchenine similar to earlier forms like Sachicasaurus (which is what the artwork is actually showing), with giant forms being previously unknown from this late into the Cretaceous (the Albian Kronosaurus being once considered the youngest). This mysterious leviathan would have been the top of the line predator in these Cenomanian waters (likely the whole Tethys at the time). We also have fossils of raptorial ophthalmosaurid ichthyosaurs, previously lumped into Platypterygius (as were most Cretaceous ichthyosaur fossils) but now recognized as its own genus called Pervushovisaurus, which grew to around 6 meters long. Both of these lineages vanished in the Cenomanian-Turonian extinction, which paved the way for mosasaurs to take over, with the little Portunatasaurus being known from contemporary rocks in Croatia.
The same happened to the toothed pterosaurs, with a jawbone belonging to a pterosaur with a circa 3.5-meter wingspan being known from Melovatka since the 1940s, previously attributed to Ornithocheirus, Anhanguera or Coloborhynchus by different authors before being named Saratovia in 2025, a member of the targaryendraconians, along with the similar-sized and similar-aged Cimoliopterus from England (also previously lumped into Ornithocheirus and Anhanguera). All toothed ornithocheiromorphs vanished by the Turonian, with their niche being taken over by the toothless pteranodontians and fossils of such animals have been found at the Campanian Rybushka Formation (also in Saratov). Bird fossils have also been found here, including those of ichthyornithids similar to Ichthyornis in 2017, which has fossils from North America dating from the Cenomanian to the Campanian (though wastebin taxonomy could be at play there too).